A northern pintail duck was also killed and three other flamingos were injured, according to the zoo.
The Rosamond Gifford Zoo says Elmer and Lima are its first same-sex parents to successfully hatch an egg, calling the penguins "exemplary in every aspect of egg care."
From elephants to otters to armadillos, all sorts of animals get a kick out stomping, rolling and eating the orange gourds.
The births will be the first for Menari, zoo officials say. She is expected to give deliver the twins sometime this winter.